Microbeam irradiation reveals cytoplasmic damage contributes to micronuclei induction Teruaki Konishi, Jun Wang, Alisa Kobayashi, Cuihua Liu, Yota Hiroyama, Taisei Mamiya, Tamon Kusumoto and Masakazu Oikawa Eur. Phys. J. Plus, 140 4 (2025) 334 Published online: 23 April 2025 DOI: 10.1140/epjp/s13360-025-06265-5